Limb Coupling and Decoupling in a Quadrupedal Walker: Can We Decipher It from a Trackway?

Deriving a gait from a trackway is a non-trivial task: Formally identical trackways are produced by different gaits of short-bodied and long-bodied animals. If the trackmaker is unknown, the gait cannot be, generally, derived from the trackway alone. I try to solve this task on an example of a fossil trackway, which is said to belong to a few-meter-long pareiasaur reptile which lived in northern Europe and became extinct more than 250 million years ago.